The Marsh Warbler - A Gloucestershire Retrospective
by Andrew Bluett
Published
in The Gloucestershire Naturalist No 18 June 2007
Introduction
The
history of the Marsh Warbler in Gloucestershire is poorly documented in spite of
it having been a noteworthy and much observed summer visitor to the county for
100 years. Few records were available from before the 1960s and the standard
reference works for the county by Witchell & Strugnell (The Fauna &
Flora of Gloucestershire, 1892), Mellersh (A Treatise on The Birds of
Gloucestershire, 1902), and even from Swaine (Birds of Gloucestershire, 1982)
are painfully brief on the subject. This is quite remarkable when compared with
other counties where it was recorded and written about in much detail, and in
light of the fact that there was no shortage of naturalists either local, or
visiting Gloucestershire, in pursuit of Marsh Warblers.
Initially
it seemed that these observers had left little or nothing to record their visits
other than a few photographs and sound recordings, but after much research a
different story has emerged. Diaries, notes and journals, eggshells in mahogany
cabinets together with the precious data cards that detail the timescale,
habitat and distribution of the Marsh Warbler have all been found and used to
construct a detailed history. Most of these records were scattered, not well
known or deliberately concealed, so that it has been largely through the kind
assistance of curators and keepers across Great Britain and the USA that data
has been unearthed from where it now resides. For that help I willingly
acknowledge and thank those people deeply.
There
is no intention here to justify or condone the actions of collectors; the taking
of eggs and skins throughout the time that it was done was illegal but the
ironic truth is that collectors’ data provided more than 80% of the
information that has made this history possible. This kind of material is too
often ignored or simply not utilised by researchers because of the stigma
attached to it, or because they do not know where to find it in spite of the
fact that museums at all levels are brimming with a hidden resource that does
little but gather dust.
The Marsh Warbler is now one of Britain’s
rarer species, and in Gloucestershire its status as “rare passage migrant”
is correct and the most apt description. Sadly, we are the poorer for that; In
the 21st century the Marsh Warbler is little more than a fond memory
for those that knew it previously, or a hoped for “tick” to the modern
observer.
An
Enigmatic Bird
The Marsh Warbler is in many ways a
mysterious creature; easily confused with its near relative the Reed Warbler
unless singing, and especially so on migration.
The question of first identification was
confused and various authors offered diverse but more or less incorrect
opinions. Christopher Swaine however wrote in Birds of Gloucestershire (1982)
that “it was not recognized as a distinct species until 1798” and whilst he
quotes no source to substantiate this, the reference must be to Bechstein and
his identification of Marsh Warblers in Germany. From
a British point of view, Eric Simms (NN71 British Warblers) mentions Blyth
separating it from the Reed Warbler here only as late as 1871 whilst S P Saville
wrote in The Zoologist of 1861 giving “Notice of the discovery and capture,
for the first time in the British Isles, of the Marsh Warbler (Calamorpha
palustris)”. Simms was clearly wrong, Saville was possibly correct, there
were certainly breeding records from 1863 (Alresford, Hants) and later in the
1860s, but there is other, albeit tenuous, evidence to suggest that Marsh
Warblers may have been recorded in this country from soon after 1840.
The most questioned and least satisfactorily
explained matter however is that of the loss of Marsh Warblers from Britain
whilst during a similar period, the population expanded far and rapidly through
mainland Europe. There is no clear evidence to explain why this might have been
and it is probably too late to discover the truth. It is difficult to carry out
such research retrospectively, so in this respect at least, the Marsh Warbler
will remain something of an enigma.
A
National & International Summary
There are sufficient records with substantive
provenance to show that breeding has been reported from more than 20 English
counties. Historically the most
significant populations were in Kent, Sussex, Somerset, Worcestershire and
Gloucestershire, the latter two accounting for more than 60% of the population
over the span of it’s tenure to date. Whilst
there is still a small population in Kent and possibly a few on Humberside, in
most regions the status of the species has been reduced to scarce, occasional,
sporadic or other equally depressing adjectives.
In the Western Palaearctic however, the Marsh
Warbler is not in the least rare. Its distribution in the breeding season covers
a vast area bounded in the south by line drawn roughly from Brittany in north
western France south eastwards to eastern Turkey, Iraq and Iran. North of that
line it is found throughout Europe in suitable habitat east to the Urals and
north to Norway, Sweden, Finland and Russia beyond the Arctic Circle as far as
latitude 67o. The population within that area is estimated to be
between 2.5 and 3.2 million pairs. The dynamics of the European population since
1900 has been of an almost uninterrupted expansion northwards reaching Sweden in
the 1920s, Finland in 1944, the Leningrad area by the 1960s, Norway in the 1970s
and the Arctic Circle by the 1980s.
Geographical
Consideration of Gloucestershire
For the purposes of this account the records
under consideration are by definition historic and in order to properly and
consistently consider Gloucestershire as it was when the Marsh Warbler’s story
began, the modern convention of vice counties is ignored and the boundaries of
“old” Gloucestershire are relied upon, when the county contained the
northern half of Bristol and district, circa 1900.
Distribution
in Gloucestershire
To Marsh Warblers, political divisions and
vice-counties were of no consequence, the truly important features were
geophysical, that is to say the wetlands and waterways along which the
populations were formerly distributed. These were continuous across and without
the county boundary. Consequently, each part of the population within
Gloucestershire was linked to those in adjacent counties. The very small numbers
of birds in the north east were essentially a part of the Evenlode, Windrush and
Cherwell population in north Oxfordshire, the Thames area birds were connected
to the Berkshire, south Oxfordshire and Wiltshire populations mainly along the
courses of the Thames and Kennet. In the south west the birds close to the Avon
were probably an extension of the Thames population and also connected to the
Somerset birds. From there it is not difficult to project a line across the
Severn estuary to the Wye and Usk, leading into the later Monmouthshire and
Herefordshire sites. The upper Severn population around Tewkesbury was linked
along the Severn into Worcestershire north to the Teme and east along the Avon
into Warwickshire.
Nowhere though was the Marsh Warbler’s
distribution fully continuous; there were significant gaps between loose
colonies or “clusters” of breeding pairs in spite of there having been at
least some apparently suitable habitat between them. There appeared to be few,
if any, differences between the areas in which there were Marsh Warblers and
those where there were none. This
is comparable with other species; the Corn Bunting shows similar tendencies even
where it is most numerous and in extremis
maintains quite isolated populations as are those on the Outer Hebrides.
South
East and South West
The earliest definite Gloucestershire record
comes in 1886 from Siddington, just to the south of Cirencester, and arises from
a nest near the Thames and Severn Canal. This is significant for being the first
record, but its location is more closely connected to the Oxfordshire/Thames
population of Marsh Warblers than anything else. The
nearest confirmed River Thames record comes from Guy Charteris, who, whilst in
pursuit of Cuckoos on 14th June 1936, found a cuckolded Marsh Warbler
nest “near Lechlade”. Whilst Lechlade itself is in Gloucestershire, reading
the data in conjunction with other notes by Charteris, reveals that this site
was in fact adjacent to the Radcott Inn at Radcott Bridge and therefore in
Oxfordshire. The hamlet of Radcott is downriver from Lechlade and 3 miles from
Gloucestershire at the nearest point. The implication is that there were more birds upstream on the
Gloucestershire Thames, but this is unproven.
A few miles north though in 1955, two or three pairs bred near
“Constant Pit” now Pit 6 in the Cotswold Water Park which is only a few
metres from the Thames Severn Canal and 4.5 km south of Siddington. More singing
males were located nearby between 1961 and 1977.
To the south west in the Bristol area, there
were several early records. The 1899 “Birds of the Bristol Region” reported
Marsh Warbler as being “a summer migrant, arriving in late May, known from
four localities in the past six years – HCP, DTP”.
The localities are not precisely identified, described only as “…the
Avon, between Bristol and Bath”. In all probability, they were in and around
the Withy beds at Saltford (Proc. from the Bristol Naturalists Society Vol IX,
1899) which were mainly south of the river, though there were Withy beds to the
north on the Gloucestershire side. Two other sites are mentioned as being
“near Avonmouth”, probably in the wet farmland and Withy beds at Hallen
Marsh where H C Playne noted that a nest had been photographed in 1898. There is
also a confirmed record of a nest at Patchway on 19th June 1909
“…in a Bean field”. H H Davies, the Bristol area recorder noted a
“passage” record at Little Stoke, near Patchway, where he was a farmer, on
15th June 1935, though he regarded the Marsh Warbler as “extinct”
in the region by 1948.
The
North East
Having found Marsh Warbler by the Thames in
Oxford in 1889/90, the Rev. W Warde-Fowler travelled to the Bernese Oberland
specifically to familiarise himself with Marsh Warbler to be better able to
identify and research them at home. His experiences there “…caused me to pay
attention to a large Osier bed in the extreme north-west corner of Oxfordshire
which was overgrown with Meadow Sweet”. On June 5th 1891 he was
passing the Osier bed when he heard a definite Marsh Warbler and wrote “It is
the Evenlode which flows past these Osier beds, dividing the county of
Oxfordshire from Gloucestershire”. He asked H C Playne to stay with him at
Kingham and together they searched there, and along the Windrush at Bourton on
the Water where the river was “bordered by gardens, meadows, orchards, Osier
beds and everything that could make a Marsh Warbler happy”. He concludes “We
will not jump to the conclusion that the Marsh Warbler is really common and has
been constantly overlooked!” On
July 15th 1891 “I (Warde-Fowler) …made one more attempt to find
the nest in company with Mr Playne who had bicycled over from Minchinhampton,
but we were again unsuccessful. The Osier bed is of recent planting; I have
myself skated on (frozen) floods in the very place it now occupies”.
The localities of the singing birds are not
pinpointed but there were Withy beds on both sides of the Evenlode around
Kingham and Bledington. The birds were almost certainly in both Oxfordshire to
the east and Gloucestershire to the west of the river. There was sufficient
belief in Warde-Fowler’s mind that more were to be found for him to make the
effort to carry out further searches along both the Evenlode and Windrush and
whilst he didn’t record the finding of a nest in Gloucestershire, he was
certain that the singing birds he discovered were breeding.
Interestingly, his companion, H C Playne, is the same individual linked
to the Avon records between Bristol and Bath.
The
Severn Vale
In the Severn Valley proper, from 1900
onwards, came an increasing number of discoveries of Marsh Warblers by the
Severn, the Sharpness and Stroudwater Canals, the Rivers Frome and Leadon, the
Coombe Hill Canal and northwards to Tewkesbury.
From south to north, a population cluster was
centred on the Saul Junction where the Stroudwater Canal (being the western end
of the Thames and Severn Canal) crossed the Gloucester and Sharpness canal to
the Severn in close proximity to the River Frome. Between and alongside the
Stroudwater Canal and River Frome there were several long and narrow withy beds
with adjacent beds of rank vegetation. Along the Sharpness canal there were
again beds of rank vegetation on the canal edge and embankments and in the
adjacent ditches, rhynes and field edges. Just to the north of the junction and
on the east side of the canal an area of ground in one of the fields at
Packthorne Farm was utilised by the canal and docks authorities for the dumping
of dredged sludge. The viscous mixture was pumped from barges over the canal
embankment into the field where it spread and settled, drying out and becoming a
fertile ground in which stray Nettle and Dock seeds germinated, thus forming a
six acre habitat colonized by Marsh Warblers that became known as the Packthorne
sludge dump.
To the west of the Sharpness canal stood an
old and semi-derelict withy bed formerly harvested by the Coles, a renowned
family of basket-makers from Quedgeley. This was leased by the RSPB from the
canal company and became the Moreton Valence Bird Sanctuary. It was monitored
for a number of years until the lease was relinquished in 1992. At its height in
1954 Peter Conder reported that the sanctuary held 9 singing males and at least
4-5 females, Bruce Campbell made similar observations a few years later. A short
way upstream on the northern edge of the cluster was yet more rank vegetation
adjacent to the Parkend Bridge which held a couple of nesting pairs.
Away from the junction there were a few more
isolated individual pairs at Frampton and on the Cambridge Arm to the south, and
possibly more down to Sharpness and Purton. To the west, Framilode held a few
pairs and there was at least one recorded nest each at Groundless Pool and
Stonebench to the north. At Eastington Marsh Warblers were found nesting on at
least two occasions and there is even a record of a nest at Nympsfield, the eggs
from which reside in the E C Stuart-Baker collection. It is unlikely that the
site was actually in or around Nympsfield proper; it was far more likely to have
been in the valley below, close to the canal.
The next clustered population to the north
was distributed around Alney Island and Walham on the western edge of Gloucester
city. At the southern edge were the withy beds at Llanthony Brickworks and rank
vegetation at Hempsted and Lower Parting where Gilroy found several nests as
early as 1906. The variety of habitat types here accommodated pairs in the withy
beds proper, the surrounding rank vegetation and along the riverside tow paths.
Alfred Thomas brought a number of collectors to this site including the Chance
brothers. The area however was not densely populated by comparison with others
and was rather looser and less reliable.
The parting of the river led then to two
distinct areas in the east and the west. On the western arm, the withy beds at
Over Bridge and the rank vegetation at the junction of the Leadon with the
Severn accounted for perhaps 10 pairs with a few more distributed along the
Leadon, the adjacent mill race and the Hereford & Gloucester Canal towards a
pair of withy beds at Lassington. It was in the Lassington withy beds that John
Walpole Bond and Colonel Richard Sparrow in company with Alfred Thomas and
Thomas Durrett found at least six pairs in 1913. From Over northwards, the river
bank alongside the road held a few pairs up to Maisemore and another area of
Withies and rank vegetation by Maisemore Weir (Staddies Pit) held more.
On the eastern arm of the river from Westgate
through Walham to Sandhurst and Ashleworth was perhaps the most productive area
of all. The key to this was a number of withy beds cultivated in the clay pits
that formerly provided the raw materials for the Severnside, Barnwood and Walham
brickworks. Around the withy beds were scrubby field corners, beds of rank
vegetation dominated by Nettles, Meadowsweet, Willowherb and Umbellifers.
Corncrake and Quail were residents of the meadows and the Marsh Warbler shared
its habitat with nine other species of Warbler.
At least one nesting pair was regularly to be
found on the riverbank at the north end of The Quay behind the slaughterhouse.
Just upstream from Westgate Bridge was the great withy bed at Pool Meadow, also
known as Shiners Perch. This was bordered on its southern edge by a timber mill
and boatyard and straddled the railway viaduct adjoining Pump House (or Black)
Bridge. This withy bed varied in quality from year to year and might hold six or
more pairs, but on occasion only two. At Walham, around the two public houses,
The Jolly Waterman and The Globe were several more withy beds. Each was named,
though the names changed over time depending on who was maintaining and
harvesting them. To the west were Cressy’s Perch and Tandy’s Pit on the site
of the former Barnwood Brickworks, to the east progressively were Robinsons
Perch (“Robbies”), Walham Perch, “The Jolly”, Longford Cement Works,
Globe Pit, and at the north end of the stretch, opposite the Upper Parting on
the site of the Tar Works (now Ronson’s Architectural Reclamation yard) was
another large withy bed, the “Nine Acres”.
To the east of Walham close to the
electricity sub-station was a large withy bed, generally known as Machine Pit,
which held six or more pairs in its hey-day. This pit is now bisected by the
Gloucester northern by-pass, either side of the by-pass, the southern end still
exists as a derelict, but obvious pool containing ancient Willows and the
northern half, cleared of its withies, is a coarse fishing pond. This site was
perceived to be dangerous to visit by those who dared venture there since it
bordered the gypsy encampment, the occupiers of which were suspicious of
strangers and rather territorial.
Upstream from Walham towards Sandhurst stood
another withy bed, the “Top Yard”, followed by Sandhurst Pit, later to
become the Gloucestershire Wildlife Trust reserve, and then another un-named
smaller pit a few hundred metres further north. Finally, opposite The Boat at
Ashleworth Quay and close by the ferry, two more withy beds held another
half-dozen pairs. A few pairs of Marsh Warbler occasionally took up residence
between the named sites and odd pairs were often on the river bank to the north
of Ashleworth.
Upriver beyond Ashleworth a few pairs were to
be found at Wainlodes, on the
Coombe Hill Canal, near Haw Bridge and on the Long (or Handkerchief) Pool below
Apperley. Following that, a long stretch of narrow withy beds and rank
vegetation extended from the White Lion at Apperley past the clay pits below
Barrow Grove to Chaceley Stock and the brickworks there. A few pairs were
occasionally to be found on the Town Street side or in the withy beds at Norton
to the east of the river, but the overall impression of this area was of
discontinuous and widely scattered small sites holding no more than one or two
pairs each. In spite of the low numbers of pairs, birds were to be found over a
longer timescale here than most other sites. The first record is of a nest found
by a Mr Porlock on the Coombe Hill Canal in June 1917, the eggs from one canal
side nest are still held in the Oxford University Museum as part of the Oliver
Vernon Aplin collection, and the nest of the last pair by the canal was chanced
upon by a Bristol entomologist in 1972. The Apperley to Chaceley stretch was one
of the easier sites to survey and study, the narrow nature of the habitats
between the river and meadows meant that pairs of Marsh Warblers positioned at
intervals along the tow path could be located by the singing males and
territorial boundaries quite readily determined.
The final clustered group of sites was that
at Tewkesbury which extended from the Lower Lode, through Severn Ham to The
Mythe at the border with Worcestershire. The Ham was a consistent site which
usually held 6 or 7 pairs of Marsh Warblers. Guy Charteris recorded finding 70
nests here over a 10 year period from 1929 and noted “…at least nine pairs
here” in 1939. Sylvia Holland counted seven singing males in June 1959.
Elsewhere in the cluster Lower Lode usually held two or three pairs and The
Mythe a similar number.
Outside the main clusters, there were
occasionally “erratic” sites, one such being the Dumbleton Bean field colony
of seven pairs found by Charteris in 1916. This was extraordinary in that
Dumbleton is a hamlet surrounded by elevated dry farmland and isolated some 9 km
from the River Avon.
As mentioned previously, there is a marked
absence of birds from extensive areas between the clusters, at least some of
which otherwise appeared to be perfectly suitable habitat. This anomaly in the
distribution of the Marsh Warbler is something that is not fully understood. It
is striking that downstream from Lower Parting there are no recorded sightings
or instances of nesting from any part of the west bank of the Severn down to
Chepstow and the Wye. Much of the ground is less attractive than the east bank,
but there were patches of suitable habitat at Minsterworth Ham, Minsterworth
proper, Rodley, Newnham, Awre and the Lydney/Aylburton Warth areas, none of
which were more than a few hundred metres across the river from key sites used
continuously for several decades.
Population
Estimates
It is difficult to be precise about how many
Marsh Warbler pairs resided in Gloucestershire. There was no truly
representative survey or count carried out until long after the decline had set
in, so that the best estimate can only be achieved by extrapolating numbers from
the available information. This is complicated by the fact that not all sites
were known, or visited, by any observers in any given year, or even any given
decade. Various conclusions have been drawn over the years; R J B Christian
listed estimates against known sites which exceeded 80 pairs, K D Pickford and
Charlie Whitfield attempted to count possible totals and arrived at “well over
60 pairs” but they considered the northern end of the vale only. Swaine
comments that “…up to ten pairs or more were to be found nesting in each of
several favoured sites. Birds continued to breed in eight or ten localities from
Purton to Tewkesbury up to the 1930s or later.” It is difficult to see just
exactly how many pairs this represents, but it is unlikely to be less than 60.
The greatest number of confirmed breeding records in any single year was 24 in
1954 but this total came from only 40% of the possible sites. It may not be very
scientific, but taking from that measure 1% equalling 0.6 nests, a figure of 60
pairs is again achieved.
The vast majority of the confirmed and
detailed data comes from egg collectors, and they readily admit that they either
could not, or did not discover all of the possible nests. This indicates that
for any given location, the confirmed and/or probable records may still be
underestimated. However, by taking each site or cluster and accumulating the
maximum numbers found there in any year, the following figures are calculated.
|
Site
or Cluster (proven & probable nest records only) |
Earliest
confirmed year |
Latest
confirmed year |
Single
Year max number Nests |
|
Saul
& Whitminster area |
1914 |
1959 |
34 |
|
Lower
Parting, Llanthony & Hempsted |
1914 |
1934 |
7 |
|
Over,
Lassington, Maisemore & The Leadon |
1913 |
1968 |
21 |
|
Walham
inc Westgate, Sandhurst & Ashleworth |
1905 |
1984 |
42 |
|
Haw
Bridge, Wainlodes and Coombe Hill |
1907 |
1972 |
9 |
|
Apperley,
Chaceley, Lower Lode & Tewkesbury |
1929 |
1977 |
23 |
|
Thames
area, CWP, Undefined and others |
1886 |
1955 |
16 |
|
|
|
|
|
|
Potential
maximum number of pairs if all sites fully occupied in any given year |
|
|
152 |
Being objective and realistic and taking into
account weather conditions, the state of the withy beds and other environmental
factors, together with the near certainty that not all sites were occupied
concurrently, a fair population estimate for the period between the 1920s and
late 1950s would probably be in the range of 65 - 85 pairs, with some
exceptional years perhaps producing up to 100 pairs.
Habitats
The habitats for the Marsh Warbler in
Gloucestershire fall into three distinct types.
The least used habitat was agricultural;
crops such as Rape, Beans and to a lesser degree, the cereals. There are only a
few definite records of pairs in crops; Christian mentioned seeing singing males
in field corners of Oats and Rape in the upper Severn in the 50s and 60s and
Charteris discovered a small colony of seven pairs in a field of Beans on the
outskirts of the village of Dumbleton in June 1916. H H Davis found a pair
nesting in a corn field adjacent to the A38 at Moreton Valence on 24th
June 1934 and there is the 1909 Patchway record from a Bean field. These are all
comparable with Stanley Lewis’s nest in Barley in Somerset and Harthan’s
Worcestershire Bean field birds in 1938, but Marsh Warblers using crops seem to
be far more common in Europe (“….in cultivated fields chiefly of all kinds
of cereals. It is particularly frequent in Rape (Brassica napus)” –
Poland, 1949, Bronislaw Ferens). In June 2006 in Belgium, the author found a
pair in a dry ditch between two fields of Barley and at another location close
to the Dutch border there were three singing males in scattered fruit trees
along a rough field edge which almost certainly had females sitting on nests in
the weedy edges of the wheat below the trees.
In a Gloucestershire context, the two more
“normal” habitats were beds of rank vegetation associated with watercourses
but in the dryer high marsh zone, and the traditional Withy beds found along the
Severn and its tributaries, the canals, and the eastern catchments of the
Thames, Windrush and Evenlode.
The rank vegetation was dominated by
Meadowsweet, Nettle, Dock, Willowherb, Umbellifers. The rest of this plant
community is filled out with grasses, phragmites, sedges, etc. The general
picture is consistently of a deep bed of vertical plants, providing a variety of
cover and resistance to the effects of wind. On the fringes there were isolated
trees, Hawthorn (Cretaeegus
monogyna), Willow (Salix spp) or Alder (Alnus glutinosa) used
as song posts.
Perhaps the most important habitat type
though was the Withy beds. On the upper Severn from Gloucester to Tewkesbury,
they were a by-product of the brick making industry. Even a cursory glance at
the Ordnance Survey maps from around 1900 shows at least seven brickworks within
sight of Gloucester Cathedral churning out the millions of bricks required to
build the expanding urban sprawl.
The sequence of excavating brick clay close
to the river-bank resulted in pits that would hold water and begin to accumulate
silt that could then be planted with Osier stakes (Salix
viminalis) at “…about a yard-and-a-half between”. The stakes would
readily root and be left to grow for 2 or 3 years before harvesting began. The
“stools” or withy stocks had the appearance when mature of being pollarded
willows in miniature, perhaps 12 – 18 inches high. The beds had a life of 20
years or more, with best production expected after 5 or 6 years, and a decline
after perhaps 15 years. The bed could then be completely cut out and re-planted
with fresh stock and the harvesting process would resume when re-established. In
spring the Withy beds would fill with Rushes, Reed Mace, Nettles, Docks, Tansy,
Iris, and Meadowsweet, the dryer banks would have some Willowherb and
Umbellifers. This under storey grew clean and free from Cleavers or Goose Grass
in most years, since it was normally cut and cleared out before the withy
harvest to aid access. What wasn’t cut would be trampled flat and the
following year’s growth was enabled to start completely afresh.
This was a microcosmic environment managed to
a high degree for the withy production and as such, it became an artificially
enhanced super-habitat hosting a denser population and a large number of nests
over an extended period of years. The decline of the Marsh Warbler from its
numerical height coincides with the decline in the production of Withies.
Garden wildlife surveys carried out primarily in search of invertebrates
have shown that one of the most important factors supporting both variety of
species and numbers of specimens is the volume of the herbaceous growth. It was
found to be even more important than variety of plants or that the plants were
native in origin. Perhaps this indicates why the Withy beds were so productive.
The clean and fresh nature of the rank under storey brought about by the
management regime was good, but the volume of the withy canopy above made them
even better. Mike Smart refers to
this “…resembling a rainforest in miniature, buzzing with insects”, R J B
Christian mentioned “….emerging from an insect infested withy bed sweating
and stinking and bitten to death by mosquitoes!”
Beyond these habitat types, many authors and
reference books list orchards among the Marsh Warbler’s nesting habitats, and
yet, within the data and information from almost 600 nest records, not a single
one came from anything remotely resembling an orchard until a very old report in
British Birds (Vol III p 157, 1909) came to light. This article recounts the
experiences of Messrs W Davies and F Coburn discovering Marsh Warblers in
Worcestershire. Davies says “I found them chiefly along hedgerows adjoining
fields of wheat and beans. There was in most cases a ditch along the hedge, in
some cases with water, in other cases dry, in all cases there was a luxuriant
growth of coarse herbage.” The following paragraph then states “Mr F Coburn
spent a day or two in the same neighbourhood, and found a pair breeding on June
13th in an old orchard ……used only for grazing cattle and as a
fowl run”. This one reference to
an Orchard seems to have become a much quoted “fact”.
It was particularly surprising that not a single record from
Gloucestershire emanated from an orchard, since several core areas of the Marsh
Warbler’s distribution were surrounded by ancient orchards and patches of
fruit trees on both banks.
Plant
Types and Nest Positioning
K D Pickford maintained that Meadowsweet was
the most favoured host plant for the Marsh Warbler’s nest. Whilst this plant
is synonymous with Marsh Warblers in most locations, he was not entirely correct
in this assertion. In Sussex, Walpole Bond found 144 nests, of which only 51
were attached to Meadowsweet. In Gloucestershire, from 184 nests in which the
plant species is identified, 45 were in Meadowsweet, 25 in Willow (Withy
stocks), 59 in Nettle, 7 in Cow Parsley and 5 in Willowherb; the remaining 43
nests being supported by a mixture of other plants including Phragmites, Dock,
Bramble and Rose, coarse Grasses, Dogwood, Figwort etc.
Recent records from Kent have the nests most often associated with
Willowherb.
Many nests were supported by a combination of
plants which together provided a balance of vertical support and lateral
stiffness. The nest being a loose structure, not unlike that of Blackcap or
Garden Warbler and hung from the vegetation by its “basket handles” was
inclined to stay relatively level under wind load, thus avoiding spillage of the
contents. This is an important factor in the Severn vale where the predominant
south-west wind meant that many nests were more prone to wind damage than those
in other areas. There is also some
correlation between nest height and nest failures by wind damage. Kelsey and
Green found that in Worcestershire the least experienced birds were inclined to
build higher and were therefore at more risk, Pickford and Christian similarly
found at Walham that several nests built higher up in the vegetation, or in a
plant monotype (especially Nettles and Docks with a relatively large surface
area) spilled at least one egg from the clutch.
Of 57 Gloucestershire nests with sufficient
data to be able to determine the heights above ground, the average was 28 inches
(710mm), the minimum 8 (200mm) and the maximum 54 inches (1370mm). 31 nests (57%) were at, or below 24 inches (600mm), only 6
(10.5%) were at or above 40 inches (1015mm).
Plant species did not seem to affect the height of the nests in the
sample, Meadowsweet and Nettle figured in both the lowest and highest 20% of the
range, the lowest two nests were in mixed Nettle and Meadowsweet, the highest
nest was in Willowherb.
The
Historical Records
The documented story of Gloucestershire’s
Marsh Warblers begins with a single nest discovered close by the Thames and
Severn Canal by a son of Henry Plummer, owner of Plummer’s Farm on the
outskirts of the village of Siddington. This discovery was made in June 1886 and
whilst the boy was ignorant of the true significance of his find, it was
recognized and reported in The Zoologist in 1887 by Herbert W Marsden. The next
confirmed record comes from a quite different source in the form of a clutch of
eggs taken from a nest “near Gloucester” in 1900, and now held as part of
the Joseph Parker Norris collection in the San Bernardino County Museum in
California. The clutch was taken, passed to a dealer (possibly Stevens’
salerooms in London) and sold on to Norris in the USA.
During the last 10 years of the 19th
century and the first 10 of the 20th century, The Rev. W
Warde-Fowler, as noted above, was making his searches in the north east of
Gloucestershire in company with Herbert C Playne, who co-incidentally was
finding them on the Bristol/Bath Avon in the south west. Playne hailed from
Minchinhampton where his father Edward and uncle Arthur were wealthy Brewers,
one a county councillor, the other a JP. Herbert was a student at University
College Oxford and all three sponsored, subscribed and contributed to Witchell
and Strugnell’s Fauna and Flora of Gloucestershire.
In 1905 the first of the collectors arrived
in the form of Norman James Gilroy, later to achieve fame, as the discoverer of
the “secret of the Greenshank”. In spite of his nefarious intent, Gilroy was
an excellent field naturalist and a pioneer in discovering and describing the
breeding cycle of Greenshanks, and in finding and recording high arctic waders
on their breeding grounds in the Pasvik Valley in arctic Norway and Finland. His
diaries are fascinating and document not only what he pursued, but many
incidental species as well as social history of the diverse places he visited.
On one of his trips to Gloucester he mentions walking from the railway station
to Walham and noting Cirl, Yellow and Reed Bunting, Wheatear, Stonechat, Redpoll
and Red-backed Shrike along the railway embankment! He was followed soon after
by Alexander Macomb Chance and his brother, Edgar, also to achieve lasting fame
with his remarkable work on the breeding cycle of the Cuckoo, most of the
research being carried out at Hartlebury Common in Worcestershire. These three
between them found many Marsh Warblers breeding in the rank vegetation and withy
beds from Frampton in the south to Ashleworth in the north, particularly at
Llanthony Weir, Hempsted and Lower Parting, Westgate, Walham and Sandhurst
between 1905 and 1918.
Cecil Stanley Meares and Douglas Hadley
Meares were introduced to Gloucester “…through the courtesy of Edgar
Chance”. These two again found many nests in the stretch of riverbank from
Llanthony to Ashleworth. 1913 brought yet another pioneer researcher and
documenter of birds with the arrival of John Walpole Bond. Bond travelled out to
the west of Gloucester to a series of withy beds at Lassington, on the banks of
the Leadon and below Lassington Wood, in company with Col. Richard Sparrow where
they found at least 6 pairs of Marsh Warblers nesting in the course of a couple
of hours.
All of these men and many others besides were
assisted and guided by one Alfred Thomas and his assistant, Thomas Durrett.
Thomas was a taxidermist with a shop at 9 College Court, between the Cathedral
precinct and Westgate Street. He went on to achieve immortality by creating most
of the major taxidermy exhibits in what is now The National Museum and Galleries
of Wales in Cardiff for the princely salary of £4.10s a week. More of his work
resides in Manchester University Museum and a pair of Gloucestershire Marsh
Warblers in a diorama constructed by him on the recommendation of Walpole Bond,
is held in the Booth Museum in Brighton. Durrett was a local postman who by dint
of spending much of his life delivering mail around Walham and Sandhurst was
able to help pinpoint likely pairs of warblers for Thomas to offer up to his
clients.
Through the 1920s to the Second World War a
small but determined series of collectors visited Gloucester with the same
purpose in mind, to procure a set or two of eggs and to study Marsh Warblers in
their habitat. There is no doubt that what Alfred Thomas taught Walpole Bond
enabled him to go on to discover the colonies in Sussex which he studied
intensively during the 1920s. From that research he was able to produce his
authoritative and very accurate writings on the species in British Birds and
“A History of the Birds of Sussex”.
Further to the north in the Tewkesbury area,
Guy Lawrence Charteris discovered 70 nests over a 10 year period from 1929 to 39
with the aid of an assistant who gloried in the unlikely name of Tom Buggins.
Charteris also discovered two quite different situations, a small colony of 7
pairs of birds in a bean field on the outskirts of Dumbleton, and a single nest
in a gravel pit on his estate at Stanway, both of these sites being out of the
norm and several kilometres from the river. Charteris’s primary interest was
the Cuckoo, and whilst he examined many nests which he refers to in his notes
and journals, he wrote very little detail unless they contained a Cuckoo’s
egg. On finding his first cuckolded nest at Severn Ham, Tewkesbury in June 1939,
he notes two comments, “At least 9 pairs of Marsh Warbler here” and later
“MW/Cuckoo No 261 - Six Marsh Warbler nests only found this year in two visits
in bad weather. Since 1929 I had examined about 70 nests here and given up hope
of a Cuckoo's egg. At last not only a Cuckoo's egg, but one that goes
extraordinarily well with the Marsh Warbler’s eggs".
By 1925 the search had partially shifted
south from Gloucester with the coming of Montagu Leighton Ridgway, a solicitor
from Lancashire who moved to practice in Herefordshire, then retired to
Gloucestershire. Ridgway took a particular interest in the Marsh Warblers around
Saul Junction, Whitminster and Parkend Bridge.
Gilroy discovered nests at Frampton in 1918, but there is little detail
attached to these and very little else of note before Ridgway’s records in
this vicinity. Ridgway was in contact with Edgar Chance and with K D Pickford
and others, but his most notable achievement was in personally finding at least
63 nests between 1925 and 1935. No other individual found more.
Collectors though, were by no means the only
visitors, Kearton had visited before 1910, T A Coward came in 1926 and much
later in the 1950s, Eric Hosking came, all of these three were in pursuit of
photographs. The sound recordist Victor C Lewis came to Walham in 1963. He
acknowledges the help afforded by Sylvia Holland of North Gloucestershire
Naturalists Society in locating a singing male and on his disc the bells of
Gloucester Cathedral are clearly audible in the background.
Coward records how he visited “a thriving
colony” describing “The situation of some of the nests we saw was
interesting, a small Withy bed, with a smoking brick kiln on one side, and the
town tip, also smoking, on the other”. This is a direct reference to the
general view of the riverbank around Walham. Far from being the pastoral scene
recorded in Frances Frith’s photographs with Poplars and water meadows leading
the eye to the stately Cathedral, it was an unmitigated industrial scene with
brickworks complete with kilns and smoking chimneys, timber yards, boat yards, a
knackers yard and slaughterhouse, the tar works and wharves separated by patches
of waste ground and at the same time all connected by a much used towpath,
constantly trodden by horses and family teams of human draught horses pulling
barges.
By the late 1930s, home grown collectors and
naturalists appeared on the scene. Alfred Thomas the procurer was replaced by
Charles Joel Whitfield, the guide to all who wanted to see Marsh Warblers and
other species on the upper Severn. Ridgway passed on the baton to Kenneth
Pickford, a local building contractor who pursued Marsh Warbler with vigour
throughout their range in the county in company with Arthur Whittaker, the
Sheffield architect and many others. Charlie Whitfield was accompanied on
searches by Dr Oliver H Wild, curator of the city museum during WW2 whose egg
collection now also resides in the USA.
The Second World War though with increased
security and restricted travel provided a period of respite for the Warblers and
enabled them to increase to the maximum population levels they were able to
achieve. Charlie Whitfield commented that “…they were quite common in the
late 1920s”, and believed that there were at least 50 pairs between the Lower
Parting and Tewkesbury in the 1940s. This in fact was probably a conservative
estimate, numbers gleaned from various sources and especially the collectors,
points to more. Wartime industry
also brought about a strange curiosity in that the nests of the Marsh Warbler
(and other species) were constructed with the aid of an alien material. The use
of “window” for blinding radar was being developed in the skies over
Worcestershire and Gloucestershire. Window (in the modern age called
“chaff”) was something akin to Christmas tinsel, small ribbons of aluminium
foil that could be dropped from pathfinder aircraft to hide following bombers
from the ground defence radar operated by the Germans. This material drifted
down to earth where the warblers collected and wove it into the fabric of their
nests. Charlie Whitfield recalled finding nests that were almost entirely
wreathed in the silver foil and stood out like beacons amongst the greenery.
After 1950 the Withy beds were beginning to
fall into decay and becoming overgrown, plastics and other materials developed
during the war had overcome the labour intensive traditional methods of making
crates and baskets. Less people and animals used the riverbank and the
undergrowth there began to change its character, it was no longer cleared or
trampled so that it became less “clean” and was more inclined to be
contaminated with Cleavers and Bindweed. Charlie Whitfield said “Never look
for a Marsh Warbler where there are Cleavers; where a dried stem of Parsley or
Nettle from last year shows above the new growth, look there!” The brickworks
were largely gone, agriculture was changing, drainage and canalisation began to
lower water levels and the ponds stagnated and progressively filled and dried
out.
Unbeknown to many, the decline had begun,
albeit only slightly, and at first slowly. Several sources noted this, in spite
of Pickford’s assertion at the Jourdain Society dinner on 9th July
1959 in Salisbury where he said “I believe that the Marsh Warbler is far more
widely distributed over the southern half of the country in suitable localities,
than the books and authorities would have us believe. Its range extends,
unbroken, save by unsuitable terrain, right through the Severn Valley, from
Berkeley in the south, to some miles above Tewkesbury in the north. There are
well known areas of colonization in Somerset and right across the country
through Oxfordshire. Hereford and Worcestershire can also claim a few pairs, as
can a dozen other counties in southern England. Through the last 30 years, in my
own experience, no variation has been made to the status of the species in my
area. We still have just as many Marsh Warbler breeding with us as were present
in my first season in 1929”.
Pickford could not have been more wrong.
Other species in the area had also begun to disappear, Red-backed Shrike, Cirl and
Corn Bunting, Corncrake and Quail were going or gone and the Woodlark would soon
follow.
During the 1950s others came upon the scene
with a different reason for seeking out Marsh Warblers. Peter Conder of the RSPB
visited the marshy withy bed in 1954 which had become the Moreton Valence Bird
Sanctuary, to the west of the Gloucester-Sharpness canal, mid-way between Saul
Junction and Parkend Bridge. He noted 9 singing males and 4-5 females; Bruce
Campbell found a nest and counted 5-6 singing males in 1959. Morgan Phillips
Price MP searched them out north of Gloucester, though seems to have written
publicly only about Chiff Chaff, Willow Warbler and Nightingale in British
Birds. Phillips Price worked the Apperley area in his researches, Christian
recalls meeting him there “…in a vast Nettle bed on the banks of the River
Severn near Tewkesbury where he was searching for nests in those halcyon days
when disturbance of a bird was not an anti-social and illegal act. He suddenly
appeared through a haze of head-high “stingers” grasping a giant sickle with
which he was laying waste to the offending urticas as if they were the very
enemies of socialism itself!” Bren Owen of the North Gloucestershire
Naturalists Society had reservations about this behaviour and noted in his
journal that he wished Price would “stay out of the undergrowth” for fear of
disturbing the birds. Christian searched them out in many sites and wrote on the
breeding biology in the Oologist’s Record. Brenig Owen, Sylvia Holland and
others in the North Gloucestershire Naturalists Society were watching and
counting. The shift towards monitoring and conservation was about to become a
reality, albeit rather too late.
In September 1968, Pickford gave an update on
the status of Marsh Warblers to the Jourdain Society at their dinner at The New
County Hotel in Gloucester, and at which the curator of the city museum, David
Dartnell, was present. He first
quoted from his earlier offering of July 1959 and then went on to say “I wish
that this statement were true today. Sad as it may be, we must accept that the
Marsh Warbler has almost, if not completely, vanished from our shores. It is
probably true to say, in retrospect, that even while I was making my generous
claim in Salisbury nine years ago, the exodus was gaining momentum. Statistics
now show that at about this period of time there was a reduction in the number
breeding on Sedgemoor and the most westerly penetrations had already been
abandoned. Each year since that time has produced a clear indication of the
drift back towards the east and in more recent times, we have watched it move
along the valley of the Severn, by the early 1960s the birds had left
Whitminster, Saul, Packthorne (sludge dump) and Moreton Valence. In 1963 there
was no nesting west or south of Gloucester and each year since has continued the
trend of evacuation”.
Christian agreed with this general picture,
his notes reveal that there were just a few pairs scattered about to the north
and west of Gloucester, but none south of the city. Phillips-Price concurred;
both commented on the paucity of occupied sites and lack of management of the
now defunct withy beds. From the late 1960s through the 70s and into the 1980s
the records of authenticated birds and breeding occurrences become ever less
frequent and more scattered. Charlie Whitfield managed to find and show a few
nests at Walham to Brenig Owen, the NGNS members found a few more around
Sandhurst, Apperley and Tewkesbury, Mike Smart found a singing bird in 1984 and
in the winter of 1984/85 John Sanders discovered a used nest at Sandhurst. There
was possibly a final breeding pair there in 1985, but to all intents and
purposes, by 1986 the story was over, just 100 years after it began.
After that time, the few records received were of isolated, transient or
passage birds discovered mainly by ringers.
The
Marsh Warbler and The Law
In terms of legal protection, most people
think of the 1954 Protection of Birds Act as being the first important
legislation. In fact the 1954 Act was a relative latecomer; there had been a
number of very much earlier pieces of legislation as far back as the Sea Birds
Preservation Act of 1869 and the Wild Birds Protection Act of 1872.
One of the key pieces of legislation however
was the Wild Birds Protection Acts 1880 to 1896. The reason for the importance
of these Acts was that they brought into play specifically local legislation or
“County” orders. For Gloucestershire these were the “Wild Birds Protection
(Administrative County of Gloucester) Orders” from 1900 onwards, designed to
allow protection to be specified at County level, but enacted under an order
issued in Whitehall. County Orders were renewed every 5 years or so.
The 1900 Order specifically mentions Marsh
Warbler in a list of 23 species along with Wood and Grasshopper Warbler, Osprey,
Nightingale, Wryneck, Woodpeckers and others for which the “…taking or
destroying of the eggs of the following species of Wild Birds is prohibited
throughout the whole of the County of Gloucester”.
A new Order was made in 1905, followed by another on 17th
October 1910, signed into being by no less an authority than Winston S.
Churchill, then Home Secretary in the Liberal Government headed by Herbert
Asquith. Another Order was to follow in 1915 and later, further protection was
offered by the Protection of Birds Acts of 1925 and 1933.
The cumulative effect of these Acts and
Orders was to make illegal all of the egg collecting activities carried on from
1900 onwards, until the 1954 Act that named the Marsh Warbler as a Schedule 1
species and introduced “special penalties”, followed by the 1981 Wildlife
and Countryside Act which also made illegal “wilful disturbance”. In spite
of the protection on offer there was no real barrier to the likes of Alfred
Thomas carrying on his procuring activities and indeed his taxidermy.
It is ironic that by the time real protection
and public awareness were on offer, it was already too late for any benefit to
accrue to the Marsh Warbler, moreover as an archetypal “little brown job” it
would hardly have attracted the protective attention reserved for the big, the
beautiful and the birds of prey.
Decline
and Loss
The question that has been asked more than
any other through my researches is that of the reasons for the loss of Marsh
Warblers, both here in Gloucestershire and in Britain as a whole.
The following comments are only a brief overview, but may shed some
light; In all honesty I fear we will never know for sure, it is almost certainly
too late to test any theory that might be proposed.
Habitat - Several authors and commentators
have somewhat glibly suggested “habitat loss” but this is at least not
wholly right. Christian expressed such an opinion in September 1967, emphasising
that whilst it was a contributing factor, “…habitat loss did not altogether
explain matters”. This view is perhaps reinforced by the fact that habitat
management was practised latterly in Worcestershire, but did little to stem the
loss there. There is little doubt
that the loss of the withy beds was significant since they appeared to be a
reservoir of higher productivity than the unmanaged environments of the rank
vegetation but more than half of the nest records came from beyond the withies,
albeit at a lower density. To all intents and purposes the general habitats
available now are no different from those in France, Belgium and the Netherlands
where Marsh Warblers continue to thrive.
Climate - Over the period of Marsh Warbler
occupation of Britain the average temperatures have changed. There was a warming
cycle that for example caused a cessation of the “frost fairs” held on the
frozen Thames in the late 18th and early 19th centuries.
The 1860-80 period was warmer, though still relatively cool in summer, and there
is a perhaps a correlation between the rise in numbers of Marsh Warblers and the
rise in average temperatures as the environment warmed slowly to the long hot
summers of the 1940s. The subsequent cooling to the 1980s before the current
rapid phase of global warming became evident is also concurrent with the decline
of the Marsh Warbler. This is a very simplified picture of a complex issue and
without detailed records for a real comparison there is no way of knowing if it
is significant. On a Eurasian-wide
scale the expansion of the Marsh Warbler’s range and numbers particularly in
the north, may be a reflection of a general warming of the continental weather
patterns. Summer rainfall varies from year to year but overall there does not
seem to have been so much change as to unduly influence the Marsh Warblers. From
time to time a bad June occurs with higher than normal rainfall, sometimes
accompanied by wind and it may have a local and periodic effect, particularly if
unseasonal flooding occurs. Christian mentions in 1955, that he and Charlie
Whitfield discovered a nest in the “Top Yard” that was suspended just inches
above flood waters in Meadowsweet.
Food Supply - The Marsh Warbler is an
insectivorous species; it is also a long distance migrant. These two factors may
hold a clue to reduced productivity. To take different species as an example of
the possible explanation, recently published information from continental
studies shows that the Pied Flycatcher is suffering from insect food sources
having shifted their life cycle forward by as much as sixteen days. The
Flycatchers are, like the Marsh Warbler, long distance migrants and are
therefore much less able to match this shift, as a result laying productivity
has fallen and nestling mortality has risen; thereby reducing the population
over time. In all probability, several later arriving species including Wood
Warbler and Redstart are suffering the same fate. The Blackcap in contrast has
reduced its migration pattern so that many of them now manage to over-winter in
Britain and Europe, which to some degree may be helping to raise the population
levels since they are able to take advantage of being able to lay earlier, and
increasing the number of second broods.
The “shallow” Gene Pool - In Britain, the
Marsh Warbler was on the outer fringe of its range. It was relatively isolated
from the European populations and with a total of perhaps no more than 250 pairs
at best the gene pool was always going to be quite limited. Kelsey and Green
suggested that there was very little if any incoming stock to bolster the
Worcestershire population. Other populations such as Walpole Bond’s Sussex
birds never produced more than 25-30 pairs and were only found over a relatively
short period of circa 20 years, so that they were in effect, a transitory
population.
An isolated and small population is always
susceptible to implosion and collapse by means of the threats from disturbance,
predation, habitat loss and climate change. Low viability and susceptibility to
health risks due to reduced genetic diversity must make this more likely.
Again, in contrast to the European birds, Kelsey and Green found that
productivity was lower in Worcestershire than in Europe. Anecdotal evidence from
several observers suggests that clutch size in Europe tends to be greater on
average with many more occurrences of 6 eggs than in the Gloucestershire (and
other British) records. There is even one record of 7 eggs in a nest near Ghent
in Belgium in 1992. Of 342 recorded Gloucestershire clutches, only 10 were of 6
eggs.
Kelsey et al make mention of the numbers of
unmated males in Worcestershire ranging from 30-40% in the mid 1980s, when the
decline elsewhere was almost complete. This suggests that whilst the males were
site faithful and returning to imprinted locations, the females were either not
surviving, or perhaps simply not returning to their birth localities. In most
species, the male holds a territory, the female approaches and decides whether
or not she will stay. If then the prospects of encountered sites and unmated
territory holding males along the route are attractive enough to stop the
progress of migrating females short of the potential destination, there will
naturally be a decline in the number of successfully breeding pairs at the
greatest distance from the wintering grounds. It is perhaps possible that an
expanding population in Europe was actually soaking up many of the available
females on their way to British sites, before they made the Channel crossing.
Retreat South to North - One of the most
interesting aspects of the Marsh Warbler’s decline and loss is that it was
almost a mirror image of the situation in Europe, where from at least 1900 there
was an inexorable expansion northwards. The British population from the 1930s
disappeared progressively from south to north. The bulk of the breeding
populations had gone from Sussex by the late 1930s, from Somerset by the 1970s,
Gloucestershire by the mid 1980s and finally from Worcestershire by the early
1990s. This situation appears to defy logic; a retreating population should be
drawing back from the outer fringes of its range towards the core population
strongholds, whereas in fact the opposite has been the case.
This also suggests that the strongest
populations were in Gloucestershire, Worcestershire and to a lesser degree
Somerset and the smaller groups in other counties were less stable, less well
established and more vulnerable to any change in their viability and the
environment. The current Marsh Warbler conservation initiative in Kent is
tempered by the fact that observers there believe that their population is
mainly dependent upon “overshoots” from the near continent and is therefore
not truly endemic and stable.
Egg (and Skin) Collecting – there is no
doubt that egg collecting in a declining (or any) species is not helpful, and
the data used in this research indicates that more than 500 nests were stripped
of their contents over a period of 80 years or so. However, these numbers equate
to less than an average of 6.5 nests per season and most of the pairs would in
all probability have laid repeat clutches. The repeats would have been less
productive (4 eggs versus 5 in a first clutch) and would probably have had a
lower survival rate but the direct loss would have at least been partially
mitigated by the repeat process. From the available data it appears that
Gloucestershire was the prime source of eggs for collectors, Worcestershire and
Somerset were relatively much less troubled, but lost their Marsh Warblers
anyway. The conclusion then must be that collecting was probably not so
significant as to have been a major contributing factor (especially when
compared with the effects of human interference on other species). It is also a
fact that other bird species in the localities and habitats occupied by Marsh
Warblers that were not collected either to the same degree, or even at all, also
disappeared from the county during the same time frame.
Summary
There is no simple conclusion to be drawn
from any of the above. The most likely explanation for the loss of the Marsh
Warbler is a combination of factors, coupled with the fact that at a maximum of
250 pairs the whole British population was vulnerable, isolated and perhaps
barely viable. Even at its height and in the most consistent locations, the
density of breeding pairs here was significantly lower than in continental
Europe. The fact that the species was able to expand northward there suggests a
level of productivity much higher than that required to merely sustain the
population. Even now, whilst there are some local fluctuations, as in Germany
and Belgium, the population in Europe generally remains consistently high and
productive with little or no change in most areas.
The
Future
So far as Britain is concerned, the future
for the Marsh Warbler is rather uncertain. The management project currently
under way in Kent has had varied success from year to year and the opinion is
that the small population there is not very stable or consistent.
The influx of Cetti’s and to a lesser
degree Savi’s Warbler from Europe has followed a similar pattern to the Marsh
Warbler of northward expansion from their formerly southern and central European
ranges. Latterly the Zitting Cisticola, (formerly Fan-tailed Warbler) has also
managed to reach the north western coast of France and southern Holland. On the
evidence of these species and bearing in mind the Marsh Warblers headlong rush
into the northern Baltic, there may be hope of a return, though it is difficult
to see why re-colonization should not already have occurred if the habitat, food
sources and climate are still suitable. The latest information is that many
continental species, particularly insects, are invading southern Britain, some
continental birds are matching this movement but many others (Icterine Warbler
for example), do not appear ever to have been inclined to make the leap of faith
and cross the channel. We may never
see the Marsh Warbler as a breeding species in Gloucestershire again, but must
always live in hope.
Acknowledgements:
I acknowledge and thank the following for historical records, help and assistance, (posthumously in many cases):
R J B Christian, C J Whitfield, K D Pickford,
N J Gilroy, J Walpole Bond, the Chance Brothers, the Meares brothers, Alfred
Thomas, G Tomkinson, G Charteris, M Phillips-Price MP, M Smart, P Duddridge, J
Sanders, M A Hope, and others for their records, notes, diaries, journals and
personal recollections.
The Curators and Keepers of museum
collections, particularly Bob McGowan at The Royal Scottish Museum, Edinburgh,
Douglas Russell at the Natural History Museum at Tring, John Campbell at
Oxfordshire Museums Service, Malgosia Novak Kemp at The Oxford University Museum
of Natural History, Alan Price at Oldham Museum, Chris Yates at Kirlees Council,
Henry McGhie at Manchester University Museum, Peter Howlett at The National
Museum and Galleries of Wales, Rene Corrado at The Western Foundation for
Vertebrate Zoology, California, USA, Dr Jean L Woods at The Delaware Museum of
Natural History, USA, Rhonda Almager at San Bernardino County Museum,
California, USA and others who have been generous and exceedingly helpful. Chris
Jones at Gwent Ornithological Society, Richard Baatsen, Gloucestershire Bird
Recorder, Ian Carle at Gloucestershire Centre for Environmental Records, and the
staff at The British Library for historical publications.
References:
The Fauna & Flora of Gloucestershire,
Witchell & Strugnell 1892
A Treatise on The Birds of Gloucestershire,
Mellersh 1902
Birds of Gloucestershire, Swaine 1982
Gloucestershire Bird Reports, various dates
Cheltenham & District Naturalists’ Society & Gloucestershire Naturalists’ Society Journals, Reports and other published material.

Map of the Severn Vale showing confirmed breeding tetrads for Marsh Warbler.

Distribution of withy beds around Alney Island

Charlie Whitfield with a Marsh Warbler’s nest


Marsh
Warbler nests
Further
Research
My research into Gloucestershire’s Marsh
Warblers continues, but becomes more difficult as the available sources and data
become more obscure and less easy to access. Any reader who may be able to
furnish information or add a little more to the story is welcome to make
contact; all contributions, whether specific or anecdotal would be valued and
appreciated and would remain confidential if necessary.
I would be particularly pleased to see or to
borrow if possible, photographs relating to any aspect of this story, or to
receive information regarding the whereabouts of manuscript notes of the late Dr
Oliver Wild, referred to by Swaine and K D Pickford but apparently now
“lost”.
Andrew Bluett
The Gloucestershire Naturalists’ Society
50 Kingsmead,
Abbeymead,
Gloucester,
GL4 5DY
© Andrew Bluett 2008